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Mary bunun yerine yurdu İskoçya'ya geri döner ve hakkı olan tahtı geri ister. Fakat İskoçya ve İngiltere'nin yönetimi, Mary’nin kuzeni olan etkileyici genç Kraliçe Elizabeth'in (Margot Robbie) yönetimi altındadır. İki genç kraliçe erkek egemen bir dünyada hem tahtta hem de aşkta rakip konumundayken bağımsızlığa karşı evlilik oyununu nasıl oynayacaklarına karar vermelidir. 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Yolcuların ve ailelerinin yaşadıklarına da yer verilen dizinin başrolünü usta oyuncu Sean Penn üstleniyor. The First’ün kadrosunda Penn’e Natascha McElhone, Lisa Gay Hamilton, Anna Jacoby-Heron, Rey Lucas, Keiko Agena, Oded Fehr ve Hannah Ware gibi isimler eşlik ediyor. 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